32:speciation&14:speciation&2:duplication&0:leaf&1:leaf&13:speciation&9:speciation&5:speciation&3:leaf&4:leaf&8:duplication&6:leaf&7:leaf&12:speciation&10:leaf&11:leaf&31:speciation&27:speciation&21:duplication&19:speciation&17:speciation&15:leaf&16:leaf&18:leaf&20:leaf&26:speciation&24:speciation&22:leaf&23:leaf&25:leaf&30:speciation&28:leaf&29:leaf